og 264 using 修饰的问题

264. Scientists believe that unlike the males of most species of moth, the male whistling moths of Nambung, Australia, call female moths to them by the use of acoustical signals, but not olfactory ones, and they attract their mates during the day, rather than at night.
(A) by the use of acoustical signals, but not olfactory ones, and they attract
(B) by the use of acoustical signals instead of using olfactory ones, and attracting
(C) by using acoustical signals, not using olfactory ones, and by attracting
(D) using acoustical signals, rather than olfactory ones, and attract
(E) using acoustical signals, but not olfactory ones, and attracting
Choice D, the best answer, is concise, maintains parallel structure, and clearly conveys the comparisons being made between the two types of moth. In A and E, the comparison between most male moths and the male whistling moth is not clear. The use of but not does not clearly convey that most other moths use olfactory signals; rather than would be preferable, as well as parallel to rather than at night. In A, the phrase by the use of is unnecessarily wordy, and the insertion of they is not required. In E, the final verb should be attract (parallel to call), not attracting (parallel to using). Choice B violates parallelism with by the use of... instead of using, as well as with call... and attracting. Choice C distorts the meaning of the original with its suggestion that male whistling moths call female moths to them both by using acoustical signals and by attracting their mates during the day. The insertion of using in not using olfactory ones is unnecessary.

我知道比较选择D,但是对D有疑惑
using acoustical signals是现在分词短语, 那么放在句中时候应该作定语修饰
前面的名词moth的,感觉总是说不通啊, 我总是觉得using前面有by比较爽
by using...表示一种动作方式, 是不是原文就是省略了by的呢?