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compound predicate

Neanderthals had a vocal tract resembling an ape's[U] and so were probably without language[/U],a shortcoming that may explain why they were supplanted by our own species.
   
    Here,I can't understand the second member being which has been underlined.Can anyone who catch it clealy give an explanation?
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那么so呢?它的作用是什么呢?

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我觉得这里主语是Neanderthals,谓语是2个并列的词:had...and were...
优秀是一种习惯。

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