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GWD SC 题不懂,求助!

Studying the fruit fly, a household nuisance but a time-honored experimental
subject, has enabled the secrets of how embryos develop to begin to be
unraveled by scientists,

A  Studying the fruit fly, a household nuisance but a time-honored experi-mental subject, has enabled the secrets of how embryos develop to begin to be unraveled by scientists.  

B.By the study of the fruit fly, a household nuisance and also a time-honored experimental subject, it was possible for the secrets of how embryos develo0p to begin to be unraveled by scientists.  

C .By studying a household nuisance but a time-honored experimental sub-ject, the fruit fly enabled scientists to begin to unravel the secrets of how embryos develop.  

D .By studying the fruit fly, a household nuisance and also a time-honored experimental subject, the secrets of how embryos develop are begin-ning to be unraveled by scientists.  

E. The study of the fruit fly, a household nuisance but a time-honored exper-imental subject, has enabled scientists to begin to unravel the secrets of how embryos develop.  
我选的A,答案是E,认为E中enable的发起者可以是subject或The study of the fruit fly。所以觉得E不对,求NN解释,为何选E!谢谢!
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E项不存在歧义吗?我觉得E项的发起者不明啊,求解答!

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还有 A中动名词 Studying 是否可以用同位语!!!!这个值得考证!

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这个解释比较牵强,因为A选项的a household nuisance but a time-honored experimental subject是做同位语的。C和D倒是存在studying的逻辑主语问题。
比较A和E,主动比被动好。

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