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OG12-70

His studies of ice-polished rocks in his Alpine
homeland, far outside the range of present-day
glaciers, led Louis Agassiz in 1837 to propose the
concept of an age in which great ice sheets had
existed in now currently temperate areas.
(A) in which great ice sheets had existed in now
currently temperate areas
(B) in which great ice sheets existed in what are
now temperate areas
(C) when great ice sheets existed where there were
areas now temperate
(D) when great ice sheets had existed in current
temperate areas
(E) when great ice sheets existed in areas now that
are temperate


答案是B,我的问题是这里的what作何理解?
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谢谢回答,但好像二者起的并不是一样的作用,不知道哪里有关于what用法的比较全面的介绍?

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now temperate areas
WHAT 就是 NOW TEMPERATE AREAS.

我们有时候讲:WHAT I DONT KNOW IS:

我猜想这两个WHAT的用法是一样的.

但官方的解释我就不知了

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