请教一个难句

wretdgdh

It was possible to demonstrate by other methods refined structural differences among neuron types ; however , proof was lacking that the quality of the impulse or its condition was influenced by these differences , which seemed instead to influence the developmental patterning of the neural circuits .(5)  
这里的instead to 如何翻译?


有可能通过其他方法来证明神经元种类间的细微的结构差异；可是，这样的证据是缺乏的，即神经冲动的性质或者状态是受这些差异所影响的，而这些差异看起来却能影响神经网络的发育模式。

himba

差异似乎反过来影响