GWD31套---2 Q33

When an active tooth in the shark’s jaws is lost or worn down, many spare teeth lie in seemingly limitless reserve, each of which are ready to slide into the appropriate position.

When an active tooth in the shark’s jaws is lost or worn down, many spare teeth lie in seemingly limitless reserve, each of which are ready to slide into the appropriate position.
Whenever an active tooth is lost or worn down, many spare teeth lie in seemingly limitless reserve in the shark’s jaws, which are each ready to slide into the appropriate position.
Many spare teeth lie in seemingly limitless reserve in the shark’s jaws, each one of which are ready to slide into the appropriate position when an active tooth is lost or worn down.
The many spare teeth lying in seemingly limitless reserve in the shark’s jaws, each one of which is ready to slide into the appropriate position whenever an active tooth is lost or worn down.

E. In the shark’s jaws, many spare teeth lie in seemingly limitless reserve, each one ready to slide into the appropriate position whenever an active tooth is lost or worn down

这个题选的是A，each of which后面谓语用are? 如果是答案错了，大家选什么？