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GWD28-Q21 语法一道~

According to entomologists, single locusts are quiet creatures, but when locusts are placed with others of their species, they become excited, change color, vibrate, and even hum.
A.
when locusts are placed with others of their species,
B.
when placing them with others of their species, then
they placing them不对
C.
locusts, when placed with others of their species,
这样出现locusts和 they 两次主语
D.
placing them with others of their species,
they placing them不对
E.
placed with others of their species, then

我选了E  请教大家,E问题在哪里?谢谢
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