对OG 语法225题的疑惑

qwrersaa

225. Minnesota is the only one of the contiguous forty-eight states that still has a sizable wolf population, and where this predator remains the archenemy of cattle and sheep.

a. that still has a sizable population, and where

b. that still has a sizable wolf population, where

c. that still has a sizable population of wolves, and where

d. where the population of wolves is still sizable;

e. where there is still a sizable population of wolves and where

答案是E，这没问题。OG解释A错是因为that引导的丛句与where引导的丛句语法上不一致。为什莫？我看过who和whose引导的丛句可以连用啊！