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两个问题 should 省略的情况 IR计入总分吗?

Combining enormous physical strength with higher intelligence, the Ns appear to have been equipped to face any obstacle the environment could put in their paths, but their relatively disappearance during the P era indicates that an ability to adapt to some environmental change led to their extinction .

这句话 led 前面是不是省略了should
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。。。。。句子是OG12里面的。应该是verb form 题

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