10. The work of mathematician Roger Penrose in the early 1970s, on the geometry of what are called aperiodic tiles, turned out to describe the architecture of a previously unknown class of crystals.

(A) what are called aperiodic tiles, turned out to describe

(B) what is called aperiodic tiles, describes

(C) aperiodic tiles, describing

(D) so-called aperiodic tiles, describe（A）

(E) aperiodic tiles, it turned out to describe

对答案没有疑问,但是想问一下A ,B 选项中的what后面动词用单数还是复数是根据called的宾语来决定吗,这样有违动词形式与主语一致啊,但这里的主语是what应该算单数还是复数呢?

谢谢..