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OG12-SC70 假如now的位置正确,E也正确吗?

His studies of ice-polished rocks in his Alpine homeland, far outside the range of present-day glaciers, led Louis Agassiz in 1837 to propose the concept of an age in which great ice sheets had existed in now currently temperate areas.

(A)  in which great ice sheets had existed in now currently temperate areas
(B)  in which great ice sheets existed in what are now temperate areas
(C)  when great ice sheets existed where there were areas now temperate
(D)  when great ice sheets had existed in current temperate areas
(E)  when great ice sheets existed in areas now that are temperate


正确选项是B,对E选项,OG解释: Now is an adverb and should be placed just after the verb are.
1.根据其解析,假如now的位置正确,E也正确吗?
2.E正确的话和B有何不同?

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我觉得,如果改成when great ice sheets existed in areas that are now temperate,应该语法上就对了----that引导定语从句,先行词就是areas。now位置也对了。但是会被认为是wordy。因为明明adj修饰名词就很简单,但是现在却用一个从句来修饰。B选项后边是宾语从句,名词性从句一种。用what引导,what在从句中作主语。in which=when。所以应该没什么意思的区别。

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thanks.

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我想问下加了B中加了what are  是什么作用

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宾语从句

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可不可以理解成 in what = where

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