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请教一道机语法题,谢谢

The work of mathematician Roger Penrose in the early 1970s, on the geometry of what are called aperiodic tiles, turned out to describe the architecture of a previously unknown class of crystals.

(A) what are called aperiodic tiles, turned out to describe

(B) what is called aperiodic tiles, describes

(C) aperiodic tiles, describing

(D) so-called aperiodic tiles, describe

(E) aperiodic tiles, it turned out to describe

work应该是单数啊,为什么选A

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what 又不指代work;what由后面的tiles可以看出是个复数~are called aperiodic tiles

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thanks  a   lot  !!!

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