求GMAT语法题一道

Honeytea

188. (26379-!-item-!-188;#058&001961)

Neuroscientists, having amassed a wealth of knowledge over the past twenty years about the brain and its development from birth to adulthood, are now drawing solid conclusions about how the human brain grows and how babies acquire language.
A. Neuroscientists, having amassed a wealth of knowledge over the past twenty years about the brain and its development from birth to adulthood, are
B. Neuroscientists, having amassed a wealth of knowledge about the brain and its development from birth to adulthood over the past twenty years, and are
C. Neuroscientists amassing a wealth of knowledge about the brain and its development from birth to adulthood over the past twenty years, and are
D. Neuroscientists have amassed a wealth of knowledge over the past twenty years about the brain and its development from birth to adulthood,
E. Neuroscientists have amassed, over the past twenty years, a wealth of knowledge about the brain and its development from birth to adulthood,

答案是A，我选的是E，不是说having+done结构不能作定语吗？