Though subject to the same wild-animal control efforts that killed off almost all the wolves in North America over the past century,  the coyote’s amazing ability of adapting to the presence of humans has enabled it to expand its range into Alaska and Central America.

1. coyote’s amazing ability of adapting to the presence of humans has enabled it to expand its
2. coyote, because of its amazing ability of adapting to the presence of humans, have been able to expand their
3. coyote, because of its amazing ability to adapt to the presence of humans, has been able to expand its
4. amazing ability of the coyote to adapt to the presence of humans have enabled it to expand the
5. amazing ability of the coyote to adapt to the presence of humans has enabled it to expand the

我选了E， 请教怎么能分辨出主语应该是coyote,而不是ability

前面半句的主语是应该是全句的主语。

是什么subject to the efforts that killed wolf呢？应该是一个动物，而不是动物的能力。中文的理解释：“虽然遭受到杀死那些狼的行为，coyote还是可以扩大自己的种群。”